Darwiniana Part 2

The hypotheses respecting the origin of species which profess to stand upon a scientific basis, and, as such, alone demand serious attention, are of two kinds. The one, the "special creation" hypothesis, presumes every species to have originated from one or more stocks, these not being the result of the modification of any other form of living matter--or arising by natural agencies--but being produced, as such, by a supernatural creative act.

The other, the so-called "trans.m.u.tation" hypothesis, considers that all existing species are the result of the modification of pre-existing species, and those of their predecessors, by agencies similar to those which at the present day produce varieties and races, and therefore in an altogether natural way; and it is a probable, though not a necessary consequence of this hypothesis, that all living beings have arisen from a single stock. With respect to the origin of this primitive stock, or stocks, the doctrine of the origin of species is obviously not necessarily concerned. The trans.m.u.tation hypothesis, for example, is perfectly consistent either with the conception of a special creation of the primitive germ, or with the supposition of its having arisen, as a modification of inorganic matter, by natural causes.

The doctrine of special creation owes its existence very largely to the supposed necessity of making science accord with the Hebrew cosmogony; but it is curious to observe that, as the doctrine is at present maintained by men of science, it is as hopelessly inconsistent with the Hebrew view as any other hypothesis.

If there be any result which has come more clearly out of geological investigation than another, it is, that the vast series of extinct animals and plants is not divisible, as it was once supposed to be, into distinct groups, separated by sharply-marked boundaries. There are no great gulfs between epochs and formations--no successive periods marked by the appearance of plants, of water animals, and of land animals, _en ma.s.se_. Every year adds to the list of links between what the older geologists supposed to be widely separated epochs: witness the crags linking the drift with older tertiaries; the Maestricht beds linking the tertiaries with the chalk; the St. Ca.s.sian beds exhibiting an abundant fauna of mixed mesozoic and palaeozoic types, in rocks of an epoch once supposed to be eminently poor in life; witness, lastly, the incessant disputes as to whether a given stratum shall be reckoned devonian or carboniferous, silurian or devonian, cambrian or silurian.

This truth is further ill.u.s.trated in a most interesting manner by the impartial and highly competent testimony of M. Pictet, from whose calculations of what percentage of the genera of animals, existing in any formation, lived during the preceding formation, it results that in no case is the proportion less than _one-third_, or 33 per cent. It is the tria.s.sic formation, or the commencement of the mesozoic epoch, which has received the smallest inheritance from preceding ages. The other formations not uncommonly exhibit 60, 80, or even 94 per cent, of genera in common with those whose remains are imbedded in their predecessor. Not only is this true, but the subdivisions of each formation exhibit new species characteristic of, and found only in, them; and, in many cases, as in the lias for example, the separate beds of these subdivisions are distinguished by well-marked and peculiar forms of life. A section, a hundred feet thick, will exhibit, at different heights, a dozen species of ammonite, none of which pa.s.ses beyond its particular zone of limestone, or clay, into the zone below it or into that above it; so that those who adopt the doctrine of special creation must be prepared to admit, that at intervals of time, corresponding with the thickness of these beds, the Creator thought fit to interfere with the natural course of events for the purpose of making a new ammonite. It is not easy to transplant oneself into the frame of mind of those who can accept such a conclusion as this, on any evidence short of absolute demonstration; and it is difficult to see what is to be gained by so doing, since, as we have said, it is obvious that such a view of the origin of living beings is utterly opposed to the Hebrew cosmogony.

Deserving no aid from the powerful arm of Bibliolatry, then, does the received form of the hypothesis of special creation derive any support from science or sound logic? a.s.suredly not much. The arguments brought forward in its favour all take one form: If species were not supernaturally created, we cannot understand the facts _x_, or _y_, or _z_; we cannot understand the structure of animals or plants, unless we suppose they were contrived for special ends; we cannot understand the structure of the eye, except by supposing it to have been made to see with; we cannot understand instincts, unless we suppose animals to have been miraculously endowed with them.

As a question of dialectics, it must be admitted that this sort of reasoning is not very formidable to those who are not to be frightened by consequences. It is an _argumentum ad ignorantiam_--take this explanation or be ignorant. But suppose we prefer to admit our ignorance rather than adopt a hypothesis at variance with all the teachings of Nature? Or, suppose for a moment we admit the explanation, and then seriously ask ourselves how much the wiser are we; what does the explanation explain? Is it any more than a grandiloquent way of announcing the fact, that we really know nothing about the matter? A phenomenon is explained when it is shown to be a case of some general law of Nature; but the supernatural interposition of the Creator can, by the nature of the case, exemplify no law, and if species have really arisen in this way, it is absurd to attempt to discuss their origin.

Or, lastly, let us ask ourselves whether any amount of evidence which the nature of our faculties permits us to attain, can justify us in a.s.serting that any phenomenon is out of the reach of natural causation. To this end it is obviously necessary that we should know all the consequences to which all possible combinations, continued through unlimited time, can give rise.

If we knew these, and found none competent to originate species, we should have good ground for denying their origin by natural causation. Till we know them, any hypothesis is better than one which involves us in such miserable presumption.

But the hypothesis of special creation is not only a mere specious mask for our ignorance; its existence in Biology marks the youth and imperfection of the science. For what is the history of every science but the history of the elimination of the notion of creative, or other interferences, with the natural order of the phaenomena which are the subject-matter of that science? When Astronomy was young "the morning stars sang together for joy," and the planets were guided in their courses by celestial hands. Now, the harmony of the stars has resolved itself into gravitation according to the inverse squares of the distances, and the orbits of the planets are deducible from the laws of the forces which allow a schoolboy's stone to break a window. The lightning was the angel of the Lord; but it has pleased Providence, in these modern times, that science should make it the humble messenger of man, and we know that every flash that s.h.i.+mmers about the horizon on a summer's evening is determined by ascertainable conditions, and that its direction and brightness might, if our knowledge of these were great enough, have been calculated.

The solvency of great mercantile companies rests on the validity of the laws which have been ascertained to govern the seeming irregularity of that human life which the moralist bewails as the most uncertain of things; plague, pestilence, and famine are admitted, by all but fools, to be the natural result of causes for the most part fully within human control, and not the unavoidable tortures inflicted by wrathful Omnipotence upon His helpless handiwork.

Harmonious order governing eternally continuous progress--the web and woof of matter and force interweaving by slow degrees, without a broken thread, that veil which lies between us and the Infinite--that universe which alone we know or can know; such is the picture which science draws of the world, and in proportion as any part of that picture is in unison with the rest, so may we feel sure that it is rightly painted. Shall Biology alone remain out of harmony with her sister sciences?

Such arguments against the hypothesis of the direct creation of species as these are plainly enough deducible from general considerations; but there are, in addition, phenomena exhibited by species themselves, and yet not so much a part of their very essence as to have required earlier mention, which are in the highest degree perplexing, if we adopt the popularly accepted hypothesis. Such are the facts of distribution in s.p.a.ce and in time; the singular phenomena brought to light by the study of development; the structural relations of species upon which our systems of cla.s.sification are founded; the great doctrines of philosophical anatomy, such as that of h.o.m.ology, or of the community of structural plan exhibited by large groups of species differing very widely in their habits and functions.

The species of animals which inhabit the sea on opposite sides of the isthmus of Panama are wholly distinct;[Footnote: Recent investigations tend to show that this statement is not strictly accurate.--1870.] the animals and plants which inhabit islands are commonly distinct from those of the neighbouring mainlands, and yet have a similarity of aspect. The mammals of the latest tertiary epoch in the Old and New Worlds belong to the same genera, or family groups, as those which now inhabit the same great geographical area. The crocodilian reptiles which existed in the earliest secondary epoch were similar in general structure to those now living, but exhibit slight differences in their vertebrae, nasal pa.s.sages, and one or two other points. The guinea-pig has teeth which are shed before it is born, and hence can never subserve the masticatory purpose for which they seem contrived, and, in like manner, the female dugong has tusks which never cut the gum. All the members of the same great group run through similar conditions in their development, and all their parts, in the adult state, are arranged according to the same plan. Man is more like a gorilla than a gorilla is like a lemur. Such are a few, taken at random, among the mult.i.tudes of similar facts which modern research has established; but when the student seeks for an explanation of them from the supporters of the received hypothesis of the origin of species, the reply he receives is, in substance, of Oriental simplicity and brevity--"Mashallah! it so pleases G.o.d!" There are different species on opposite sides of the isthmus of Panama, because they were created different on the two sides. The pliocene mammals are like the existing ones, because such was the plan of creation; and we find rudimental organs and similarity of plan, because it has pleased the Creator to set before Himself a "divine exemplar or archetype,"

and to copy it in His works; and somewhat ill, those who hold this view imply, in some of them. That such verbal hocus-pocus should be received as science will one day be regarded as evidence of the low state of intelligence in the nineteenth century, just as we amuse ourselves with the phraseology about Nature's abhorrence of a vacuum, wherewith Torricellis compatriots were satisfied to explain the rise of water in a pump. And be it recollected that this sort of satisfaction works not only negative but positive ill, by discouraging inquiry, and so depriving man of the usufruct of one of the most fertile fields of his great patrimony, Nature.

The objections to the doctrine of the origin of species by special creation which have been detailed, must have occurred, with more or less force, to the mind of every one who has seriously and independently considered the subject. It is therefore no wonder that, from time to time, this hypothesis should have been met by counter hypotheses, all as well, and some better founded than itself; and it is curious to remark that the inventors of the opposing views seem to have been led into them as much by their knowledge of geology, as by their acquaintance with biology. In fact, when the mind has once admitted the conception of the gradual production of the present physical state of our globe, by natural causes operating through long ages of time, it will be little disposed to allow that living beings have made their appearance in another way, and the speculations of De Maillet and his successors are the natural complement of Scilla's demonstration of the true nature of fossils.

A contemporary of Newton and of Leibnitz, sharing therefore in the intellectual activity of the remarkable age which witnessed the birth of modern physical science, Benoit de Maillet spent a long life as a consular agent of the French Government in various Mediterranean ports. For sixteen years, in fact, he held the office of Consul-General in Egypt, and the wonderful phenomena offered by the valley of the Nile appear to have strongly impressed his mind, to have directed his attention to all facts of a similar order which came within his observation, and to have led him to speculate on the origin of the present condition of our globe and of its inhabitants. But, with all his ardour for science, De Maillet seems to have hesitated to publish views which, notwithstanding the ingenious attempts to reconcile them with the Hebrew hypothesis contained in the preface to "Telliamed," were hardly likely to be received with favour by his contemporaries.

But a short time had elapsed since more than one of the great anatomists and physicists of the Italian school had paid dearly for their endeavours to dissipate some of the prevalent errors; and their ill.u.s.trious pupil, Harvey, the founder of modern physiology, had not fared so well, in a country less oppressed by the benumbing influences of theology, as to tempt any man to follow his example. Probably not uninfluenced by these considerations, his Catholic majesty's Consul-General for Egypt kept his theories to himself throughout a long life, for "Telliamed," the only scientific work which is known to have proceeded from his pen, was not printed till 1735, when its author had reached the ripe age of seventy-nine; and though De Maillet lived three years longer, his book was not given to the world before 1748. Even then it was anonymous to those who were not in the secret of the anagrammatic character of its t.i.tle; and the preface and dedication are so worded as, in case of necessity, to give the printer a fair chance of falling back on the excuse that the work was intended for a mere _jeu d'esprit_.

The speculations of the suppositious Indian sage, though quite as sound as those of many a "Mosaic Geology," which sells exceedingly well, have no great value if we consider them by the light of modern science. The waters are supposed to have originally covered the whole globe; to have deposited the rocky ma.s.ses which compose its mountains by processes comparable to those which are now forming mud, sand, and s.h.i.+ngle; and then to have gradually lowered their level, leaving the spoils of their animal and vegetable inhabitants embedded in the strata. As the dry land appeared, certain of the aquatic animals are supposed to have taken to it, and to have become gradually adapted to terrestrial and aerial modes of existence.

But if we regard the general tenor and style of the reasoning in relation to the state of knowledge of the day, two circ.u.mstances appear very well worthy of remark. The first, that De Maillet had a notion of the modifiability of living forms (though without any precise information on the subject), and how such modifiability might account for the origin of species; the second, that he very clearly apprehended the great modern geological doctrine, so strongly insisted upon by Hutton, and so ably and comprehensively expounded by Lyell, that we must look to existing causes for the explanation of past geological events. Indeed, the following pa.s.sage of the preface, in which De Maillet is supposed to speak of the Indian philosopher Telliamed, his _alter ego,_ might have been written by the most philosophical uniformitarian of the present day:--

"Ce qu'il y a d'etonnant, est que pour arriver a ces connaissances il semble avoir perverti l'ordre naturel, puisqu'au lieu de s'attacher d'abord a rechercher l'origine de notre globe il a commence par travailler a s'instruire de la nature. Mais a l'entendre, ce renvers.e.m.e.nt de l'ordre a ete pour lui l'effet d'un genie favorable qui l'a conduit pas a pas et comme par la main aux decouvertes les plus sublimes. C'est en decomposant la substance de ce globe par tine anatomie exacte de toutes ses parties qu'il a premierement appris de quelles matieres il etait compose et quels arrangemens ces memes matieres observaient entre elles. Ces lumieres jointes a l'esprit de comparaison toujours necessaire a quiconque entreprend de percer les voiles dont la nature aime a se cacher, ont servi de guide a notre philosophe pour parvenir a des connoissances plus interessantes. Par la matiere et l'arrangement de ces compositions il pretend avoir reconnu quelle est la veritable origine de ce globe que nous habitons, comment et par qui il a ete forme."-Pp. xix. xx.

But De Maillet was before his age, and as could hardly fail to happen to one who speculated on a zoological and botanical question before Linnaeus, and on a physiological problem before Haller, he fell into great errors here and there; and hence, perhaps, the general neglect of his work.

Robinet's speculations are rather behind, than in advance of, those of De Maillet; and though Linnaeus may have played with the hypothesis of trans.m.u.tation, it obtained no serious support until Lamarck adopted it, and advocated it with great ability in his "Philosophie Zoologique."

Impelled towards the hypothesis of the trans.m.u.tation of species, partly by his general cosmological and geological views; partly by the conception of a graduated, though irregularly branching, scale of being, which had arisen out of his profound study of plants and of the lower forms of animal life, Lamarck, whose general line of thought often closely resembles that of De Maillet, made a great advance upon the crude and merely speculative manner in which that writer deals with the question of the origin of living beings, by endeavouring to find physical causes competent to effect that change of one species into another, which De Maillet had only supposed to occur. And Lamarck conceived that he had found in Nature such causes, amply sufficient for the purpose in view. It is a physiological fact, he says, that organs are increased in size by action, atrophied by inaction; it is another physiological fact that modifications produced are transmissible to offspring. Change the actions of an animal, therefore, and you will change its structure, by increasing the development of the parts newly brought into use and by the diminution of those less used; but by altering the circ.u.mstances which surround it you will alter its actions, and hence, in the long run, change of circ.u.mstance must produce change of organisation.

All the species of animals, therefore, are, in Lamarck's view, the result of the indirect action of changes of circ.u.mstance, upon those primitive germs which he considered to have originally arisen, by spontaneous generation, within the waters of the globe. It is curious, however, that Lamarck should insist so strongly [Footnote: See _Phil. Zoologique_, vol. i. p. 222. et seq.] as he has done, that circ.u.mstances never in any degree directly modify the form or the organisation of animals, but only operate by changing their wants and consequently their actions; for he thereby brings upon himself the obvious question, How, then, do plants, which cannot be said to have wants or actions, become modified? To this he replies, that they are modified by the changes in their nutritive processes, which are effected by changing circ.u.mstances; and it does not seem to have occurred to him that such changes might be as well supposed to take place among animals.

When we have said that Lamarck felt that mere speculation was not the way to arrive at the origin of species, but that it was necessary, in order to the establishment of any sound theory on the subject, to discover by observation or otherwise, some _vera causa_, competent to give rise to them; that he affirmed the true order of cla.s.sification to coincide with the order of their development one from another; that he insisted on the necessity of allowing sufficient time, very strongly; and that all the varieties of instinct and reason were traced back by him to the same cause as that which has given rise to species, we have enumerated his chief contributions to the advance of the question. On the other hand, from his ignorance of any power in Nature competent to modify the structure of animals, except the development of parts, or atrophy of them, in consequence of a change of needs, Lamarck was led to attach infinitely greater weight than it deserves to this agency, and the absurdities into which he was led have met with deserved condemnation. Of the struggle for existence, on which, as we shall see, Mr. Darwin lays such great stress, he had no conception; indeed, he doubts whether there really are such things as extinct species, unless they be such large animals as may have met their death at the hands of man; and so little does he dream of there being any other destructive causes at work, that, in discussing the possible existence of fossil sh.e.l.ls, he asks, "Pourquoi d'ailleurs seroient-ils perdues des que l'homme n'a pu operer leur destruction?" ("Phil. Zool.,"

vol. i. p. 77.) Of the influence of selection Lamarck has as little notion, and he makes no use of the wonderful phenomena which are exhibited by domesticated animals, and ill.u.s.trate its powers. The vast influence of Cuvier was employed against the Lamarckian views, and, as the untenability of some of his conclusions was easily shown, his doctrines sank under the opprobrium of scientific, as well as of theological, heterodoxy. Nor have the efforts made of late years to revive them tended to re-establish their credit in the minds of sound thinkers acquainted with the facts of the case; indeed it may be doubted whether Lamarck has not suffered more from his friends than from his foes.

Two years ago, in fact, though we venture to question if even the strongest supporters of the special creation hypothesis had not, now and then, an uneasy consciousness that all was not right, their position seemed more impregnable than ever, if not by its own inherent strength, at any rate by the obvious failure of all the attempts which had been made to carry it. On the other hand, however much the few, who thought deeply on the question of species, might be repelled by the generally received dogmas, they saw no way of escaping from them save by the adoption of suppositions so little justified by experiment or by observation as to be at least equally distasteful.

The choice lay between two absurdities and a middle condition of uneasy scepticism; which last, however unpleasant and unsatisfactory, was obviously the only justifiable state of mind under the circ.u.mstances.

Such being the general ferment in the minds of naturalists, it is no wonder that they mustered strong in the rooms of the Linnaean Society, on the 1st of July of the year 1858, to hear two papers by authors living on opposite sides of the globe, working out their results independently, and yet professing to have discovered one and the same solution of all the problems connected with species. The one of these authors was an able naturalist, Mr. Wallace, who had been employed for some years in studying the productions of the islands of the Indian Archipelago, and who had forwarded a memoir embodying his views to Mr. Darwin, for communication to the Linnaean Society. On perusing the essay, Mr. Darwin was not a little surprised to find that it embodied some of the leading ideas of a great work which he had been preparing for twenty years, and parts of which, containing a development of the very same views, had been perused by his private friends fifteen or sixteen years before. Perplexed in what manner to do full justice both to his friend and to himself, Mr. Darwin placed the matter in the hands of Dr. Hooker and Sir Charles Lyell, by whose advice he communicated a brief abstract of his own views to the Linnaean Society, at the same time that Mr. Wallace's paper was read. Of that abstract, the work on the "Origin of Species" is an enlargement; but a complete statement of Mr. Darwin's doctrine is looked for in the large and well-ill.u.s.trated work which he is said to be preparing for publication.

The Darwinian hypothesis has the merit of being eminently simple and comprehensible in principle, and its essential positions may be stated in a very few words: all species have been produced by the development of varieties from common stocks; by the conversion of these, first into permanent races and then into new species, by the process of _natural selection_, which process is essentially identical with that artificial selection by which man has originated the races of domestic animals--the _struggle for existence_ taking the place of man, and exerting, in the case of natural selection, that selective action which he performs in artificial selection.

The evidence brought forward by Mr. Darwin in support of his hypothesis is of three kinds. First, he endeavours to prove that species may be originated by selection; secondly, he attempts to show that natural causes are competent to exert selection; and thirdly, he tries to prove that the most remarkable and apparently anomalous phaenomena exhibited by the distribution, development, and mutual relations of species, can be shown to be deducible from the general doctrine of their origin, which he propounds, combined with the known facts of geological change; and that, even if all these phaenomena are not at present explicable by it, none are necessarily inconsistent with it.

There cannot be a doubt that the method of inquiry which Mr. Darwin has adopted is not only rigorously in accordance with the canons of scientific logic, but that it is the only adequate method. Critics exclusively trained in cla.s.sics or in mathematics, who have never determined a scientific fact in their lives by induction from experiment or observation, prate learnedly about Mr. Darwin's method, which is not inductive enough, not Baconian enough, forsooth, for them. But even if practical acquaintance with the process of scientific investigation is denied them, they may learn, by the perusal of Mr. Mill's admirable chapter "On the Deductive Method," that there are mult.i.tudes of scientific inquiries in which the method of pure induction helps the investigator but a very little way.

"The mode of investigation," says Mr. Mill, "which, from the proved inapplicability of direct methods of observation and experiment, remains to us as the main source of the knowledge we possess, or can acquire, respecting the conditions and laws of recurrence of the more complex phaenomena, is called, in its most general expression, the deductive method, and consists of three operations: the first, one of direct induction; the second, of ratiocination; and the third, of verification."

Now, the conditions which have determined the existence of species are not only exceedingly complex, but, so far as the great majority of them are concerned, are necessarily beyond our cognisance. But what Mr. Darwin has attempted to do is in exact accordance with the rule laid down by Mr. Mill; he has endeavoured to determine certain great facts inductively, by observation and experiment; he has then reasoned from the data thus furnished; and lastly, he has tested the validity of his ratiocination by comparing his deductions with the observed facts of Nature. Inductively, Mr. Darwin endeavours to prove that species arise in a given way.

Deductively, he desires to show that, if they arise in that way, the facts of distribution, development, cla.s.sification, &c., may be accounted for, _i.e._ may be deduced from their mode of origin, combined with admitted changes in physical geography and climate, during an indefinite period. And this explanation, or coincidence of observed with deduced facts, is, so far as it extends, a verification of the Darwinian view.

There is no fault to be found with Mr. Darwin's method, then; but it is another question whether he has fulfilled all the conditions imposed by that method. Is it satisfactorily proved, in fact, that species may be originated by selection? that there is such a thing as natural selection?

that none of the phaenomena exhibited by species are inconsistent with the origin of species in this way? If these questions can be answered in the affirmative, Mr. Darwin's view steps out of the rank of hypotheses into those of proved theories; but, so long as the evidence at present adduced falls short of enforcing that affirmation, so long, to our minds, must the new doctrine be content to remain among the former--an extremely valuable, and in the highest degree probable, doctrine, indeed the only extant hypothesis which is worth anything in a scientific point of view; but still a hypothesis, and not yet the theory of species.

After much consideration, and with a.s.suredly no bias against Mr. Darwin's views, it is our clear conviction that, as the evidence stands, it is not absolutely proven that a group of animals, having all the characters exhibited by species in Nature, has ever been originated by selection, whether artificial or natural. Groups having the morphological character of species--distinct and permanent races in fact--have been so produced over and over again; but there is no positive evidence, at present, that any group of animals has, by variation and selective breeding, given rise to another group which was, even in the least degree, infertile with the first. Mr. Darwin is perfectly aware of this weak point, and brings forward a mult.i.tude of ingenious and important arguments to diminish the force of the objection. We admit the value of these arguments to their fullest extent; nay, we will go so far as to express our belief that experiments, conducted by a skilful physiologist, would very probably obtain the desired production of mutually more or less infertile breeds from a common stock, in a comparatively few years; but still, as the case stands at present, this "little rift within the lute" is not to be disguised nor overlooked.

In the remainder of Mr. Darwin's argument our own private ingenuity has not hitherto enabled us to pick holes of any great importance; and judging by what we hear and read, other adventurers in the same field do not seem to have been much more fortunate. It has been urged, for instance, that in his chapters on the struggle for existence and on natural selection, Mr. Darwin does not so much prove that natural selection does occur, as that it must occur; but, in fact, no other sort of demonstration is attainable. A race does not attract our attention in Nature until it has, in all probability, existed for a considerable time, and then it is too late to inquire into the conditions of its origin. Again, it is said that there is no real a.n.a.logy between the selection which takes place under domestication, by human influence, and any operation which can be effected by Nature, for man interferes intelligently. Reduced to its elements, this argument implies that an effect produced with trouble by an intelligent agent must, _a fortiori,_ be more troublesome, if not impossible, to an unintelligent agent. Even putting aside the question whether Nature, acting as she does according to definite and invariable laws, can be rightly called an unintelligent agent, such a position as this is wholly untenable. Mix salt and sand, and it shall puzzle the wisest of men, with his mere natural appliances, to separate all the grains of sand from all the grains of salt; but a shower of rain will effect the same object in ten minutes. And so, while man may find it tax all his intelligence to separate any variety which arises, and to breed selectively from it, the destructive agencies incessantly at work in Nature, if they find one variety to be more soluble in circ.u.mstances than the other, will inevitably, in the long run, eliminate it.

A frequent and a just objection to the Lamarckian hypothesis of the trans.m.u.tation of species is based upon the absence of transitional forms between many species. But against the Darwinian hypothesis this argument has no force. Indeed, one of the most valuable and suggestive parts of Mr.

Darwin's work is that in which he proves, that the frequent absence of transitions is a necessary consequence of his doctrine, and that the stock whence two or more species have sprung, need in no respect be intermediate between these species. If any two species have arisen from a common stock in the same way as the carrier and the pouter, say, have arisen from the rock-pigeon, then the common stock of these two species need be no more intermediate between the two than the rock-pigeon is between the carrier and pouter. Clearly appreciate the force of this a.n.a.logy, and all the arguments against the origin of species by selection, based on the absence of transitional forms, fall to the ground. And Mr. Darwin's position might, we think, have been even stronger than it is if he had not embarra.s.sed himself with the aphorism, "_Natura non facit saltum_," which turns up so often in his pages. We believe, as we have said above, that Nature does make jumps now and then, and a recognition of the fact is of no small importance in disposing of many minor objections to the doctrine of trans.m.u.tation.

But we must pause. The discussion of Mr. Darwin's arguments in detail would lead us far beyond the limits within which we proposed, at starting, to confine this article. Our object has been attained if we have given an intelligible, however brief, account of the established facts connected with species, and of the relation of the explanation of those facts offered by Mr. Darwin to the theoretical views held by his predecessors and his contemporaries, and, above all, to the requirements of scientific logic. We have ventured to point out that it does not, as yet, satisfy all those requirements; but we do not hesitate to a.s.sert that it is as superior to any preceding or contemporary hypothesis, in the extent of observational and experimental basis on which it rests, in its rigorously scientific method, and in its power of explaining biological phenomena, as was the hypothesis of Copernicus to the speculations of Ptolemy. But the planetary orbits turned out to be not quite circular after all, and, grand as was the service Copernicus rendered to science, Kepler and Newton had to come after him. What if the orbit of Darwinism should be a little too circular? What if species should offer residual phaenomena, here and there, not explicable by natural selection? Twenty years hence naturalists may be in a position to say whether this is, or is not, the case; but in either event they will owe the author of "The Origin of Species" an immense debt of grat.i.tude. We should leave a very wrong impression on the reader's mind if we permitted him to suppose that the value of that work depends wholly on the ultimate justification of the theoretical views which it contains. On the contrary, if they were disproved to-morrow, the book would still be the best of its kind--the most compendious statement of well-sifted facts bearing on the doctrine of species that has ever appeared. The chapters on Variation, on the Struggle for Existence, on Instinct, on Hybridism, on the Imperfection of the Geological Record, on Geographical Distribution, have not only no equals, but, so far as our knowledge goes, no compet.i.tors, within the range of biological literature. And viewed as a whole, we do not believe that, since the publication of Von Baer's "Researches on Development," thirty years ago, any work has appeared calculated to exert so large an influence, not only on the future of Biology, but in extending the domination of Science over regions of thought into which she has, as yet, hardly penetrated.

III

CRITICISMS ON "THE ORIGIN OF SPECIES"

[1864]

1. UEBER DIE DARWIN'SCHE SCHoPFUNGSTHEORIE; EIN VORTRAG, Von A. KoLLIKER.

Leipzig, 1864.

2. EXAMINATION DU LIVRE DE M. DARWIN SUR L'ORIGINE DES ESPeCES. Par P.

FLOURENS. Paris, 1864.

In the course of the present year several foreign commentaries upon Mr.

Darwin's great work have made their appearance. Those who have perused that remarkable chapter of the "Antiquity of Man," in which Sir Charles Lyell draws a parallel between the development of species and that of languages, will be glad to hear that one of the most eminent philologers of Germany, Professor Schleicher, has, independently, published a most instructive and philosophical pamphlet (an excellent notice of which is to be found in the _Reader_, for February 27th of this year) supporting similar views with all the weight of his special knowledge and established authority as a linguist. Professor Haeckel, to whom Schleicher addresses himself, previously took occasion, in his splendid monograph on the _Radiolaria_,[Footnote: _Die Radiolarien: eine Monographie_, p.

231.] to express his high appreciation of, and general concordance with, Mr. Darwin's views.

But the most elaborate criticisms of the "Origin of Species" which have appeared are two works of very widely different merit, the one by Professor Kolliker, the well-known anatomist and histologist of Wurzburg; the other by M. Flourens, Perpetual Secretary of the French Academy of Sciences.

Professor Kolliker's critical essay "Upon the Darwinian Theory" is, like all that proceeds from the pen of that thoughtful and accomplished writer, worthy of the most careful consideration. It comprises a brief but clear sketch of Darwin's views, followed by an enumeration of the leading difficulties in the way of their acceptance; difficulties which would appear to be insurmountable to Professor Kolliker, inasmuch as he proposes to replace Mr. Darwin's Theory by one which he terms the "Theory of Heterogeneous Generation." We shall proceed to consider first the destructive, and secondly, the constructive portion of the essay.