Creative Evolution Part 5

Now, it seems to us most probable that the animal cell and the vegetable cell are derived from a common stock, and that the first living organisms oscillated between the vegetable and animal form, partic.i.p.ating in both at once. Indeed, we have just seen that the characteristic tendencies of the evolution of the two kingdoms, although divergent, coexist even now, both in the plant and in the animal. The proportion alone differs. Ordinarily, one of the two tendencies covers or crushes down the other, but in exceptional circ.u.mstances the suppressed one starts up and regains the place it had lost. The mobility and consciousness of the vegetable cell are not so sound asleep that they cannot rouse themselves when circ.u.mstances permit or demand it; and, on the other hand, the evolution of the animal kingdom has always been r.e.t.a.r.ded, or stopped, or dragged back, by the tendency it has kept toward the vegetative life. However full, however overflowing the activity of an animal species may appear, torpor and unconsciousness are always lying in wait for it. It keeps up its role only by effort, at the price of fatigue. Along the route on which the animal has evolved, there have been numberless shortcomings and cases of decay, generally a.s.sociated with parasitic habits; they are so many shuntings on to the vegetative life. Thus, everything bears out the belief that vegetable and animal are descended from a common ancestor which united the tendencies of both in a rudimentary state.

But the two tendencies mutually implied in this rudimentary form became dissociated as they grew. Hence the world of plants with its fixity and insensibility, hence the animals with their mobility and consciousness.

There is no need, in order to explain this dividing into two, to bring in any mysterious force. It is enough to point out that the living being leans naturally toward what is most convenient to it, and that vegetables and animals have chosen two different kinds of convenience in the way of procuring the carbon and nitrogen they need. Vegetables continually and mechanically draw these elements from an environment that continually provides it. Animals, by action that is discontinuous, concentrated in certain moments, and conscious, go to find these bodies in organisms that have already fixed them. They are two different ways of being industrious, or perhaps we may prefer to say, of being idle.

For this very reason we doubt whether nervous elements, however rudimentary, will ever be found in the plant. What corresponds in it to the directing will of the animal is, we believe, the direction in which it bends the energy of the solar radiation when it uses it to break the connection of the carbon with the oxygen in carbonic acid. What corresponds in it to the sensibility of the animal is the impressionability, quite of its kind, of its chlorophyl light. Now, a nervous system being pre-eminently a mechanism which serves as intermediary between sensations and volitions, the true "nervous system"

of the plant seems to be the mechanism or rather chemicism _sui generis_ which serves as intermediary between the impressionability of its chlorophyl to light and the producing of starch: which amounts to saying that the plant can have no nervous elements, and that _the same impetus that has led the animal to give itself nerves and nerve centres must have ended, in the plant, in the chlorophyllian function_.[55]

This first glance over the organized world will enable us to ascertain more precisely what unites the two kingdoms, and also what separates them.

Suppose, as we suggested in the preceding chapter, that at the root of life there is an effort to engraft on to the necessity of physical forces the largest possible amount of _indetermination_. This effort cannot result in the creation of energy, or, if it does, the quant.i.ty created does not belong to the order of magnitude apprehended by our senses and instruments of measurement, our experience and science. All that the effort can do, then, is to make the best of a pre-existing energy which it finds at its disposal. Now, it finds only one way of succeeding in this, namely, to secure such an acc.u.mulation of potential energy from matter, that it can get, at any moment, the amount of work it needs for its action, simply by pulling a trigger. The effort itself possesses only that power of releasing. But the work of releasing, although always the same and always smaller than any given quant.i.ty, will be the more effective the heavier the weight it makes fall and the greater the height--or, in other words, the greater the sum of potential energy acc.u.mulated and disposable. As a matter of fact, the princ.i.p.al source of energy usable on the surface of our planet is the sun. So the problem was this: to obtain from the sun that it should partially and provisionally suspend, here and there, on the surface of the earth, its continual outpour of usable energy, and store a certain quant.i.ty of it, in the form of unused energy, in appropriate reservoirs, whence it could be drawn at the desired moment, at the desired spot, in the desired direction. The substances forming the food of animals are just such reservoirs. Made of very complex molecules holding a considerable amount of chemical energy in the potential state, they are like explosives which only need a spark to set free the energy stored within them. Now, it is probable that life tended at the beginning to compa.s.s at one and the same time both the manufacture of the explosive and the explosion by which it is utilized. In this case, the same organism that had directly stored the energy of the solar radiation would have expended it in free movements in s.p.a.ce. And for that reason we must presume that the first living beings sought on the one hand to acc.u.mulate, without ceasing, energy borrowed from the sun, and on the other hand to expend it, in a discontinuous and explosive way, in movements of locomotion. Even to-day, perhaps, a chlorophyl-bearing Infusorian such as the Euglena may symbolize this primordial tendency of life, though in a mean form, incapable of evolving. Is the divergent development of the two kingdoms related to what one may call the oblivion of each kingdom as regards one of the two halves of the programme? Or rather, which is more likely, was the very nature of the matter, that life found confronting it on our planet, opposed to the possibility of the two tendencies evolving very far together in the same organism? What is certain is that the vegetable has trended princ.i.p.ally in the first direction and the animal in the second. But if, from the very first, in making the explosive, nature had for object the explosion, then it is the evolution of the animal, rather than that of the vegetable, that indicates, on the whole, the fundamental direction of life.

The "harmony" of the two kingdoms, the complementary characters they display, might then be due to the fact that they develop two tendencies which at first were fused in one. The more the single original tendency grows, the harder it finds it to keep united in the same living being those two elements which in the rudimentary state implied each other.

Hence a parting in two, hence two divergent evolutions; hence also two series of characters opposed in certain points, complementary in others, but, whether opposed or complementary, always preserving an appearance of kins.h.i.+p. While the animal evolved, not without accidents along the way, toward a freer and freer expenditure of discontinuous energy, the plant perfected rather its system of acc.u.mulation without moving. We shall not dwell on this second point. Suffice it to say that the plant must have been greatly benefited, in its turn, by a new division, a.n.a.logous to that between plants and animals. While the primitive vegetable cell had to fix by itself both its carbon and its nitrogen, it became able almost to give up the second of these two functions as soon as microscopic vegetables came forward which leaned in this direction exclusively, and even specialized diversely in this still complicated business. The microbes that fix the nitrogen of the air and those which convert the ammoniacal compounds into nitrous ones, and these again into nitrates, have, by the same splitting up of a tendency primitively one, rendered to the whole vegetable world the same kind of service as the vegetables in general have rendered to animals. If a special kingdom were to be made for these microscopic vegetables, it might be said that in the microbes of the soil, the vegetables and the animals, we have before us the _a.n.a.lysis_, carried out by the matter that life found at its disposal on our planet, of all that life contained, at the outset, in a state of reciprocal implication. Is this, properly speaking, a "division of labor"? These words do not give the exact idea of evolution, such as we conceive it. Wherever there is division of labor, there is _a.s.sociation_ and also _convergence_ of effort. Now, the evolution we are speaking of is never achieved by means of a.s.sociation, but by _dissociation_; it never tends toward convergence, but toward _divergence_ of efforts. The harmony between terms that are mutually complementary in certain points is not, in our opinion, produced, in course of progress, by a reciprocal adaptation; on the contrary, it is complete only at the start. It arises from an original ident.i.ty, from the fact that the evolutionary process, splaying out like a sheaf, sunders, in proportion to their simultaneous growth, terms which at first completed each other so well that they coalesced.

Now, the elements into which a tendency splits up are far from possessing the same importance, or, above all, the same power to evolve.

We have just distinguished three different kingdoms, if one may so express it, in the organized world. While the first comprises only microorganisms which have remained in the rudimentary state, animals and vegetables have taken their flight toward very lofty fortunes. Such, indeed, is generally the case when a tendency divides. Among the divergent developments to which it gives rise, some go on indefinitely, others come more or less quickly to the end of their tether. These latter do not issue directly from the primitive tendency, but from one of the elements into which it has divided; they are residual developments made and left behind on the way by some truly elementary tendency which continues to evolve. Now, these truly elementary tendencies, we think, bear a mark by which they may be recognized.

This mark is like a trace, still visible in each, of what was in the original tendency of which they represent the elementary directions. The elements of a tendency are not like objects set beside each other in s.p.a.ce and mutually exclusive, but rather like psychic states, each of which, although it be itself to begin with, yet partakes of others, and so virtually includes in itself the whole personality to which it belongs. There is no real manifestation of life, we said, that does not show us, in a rudimentary or latent state, the characters of other manifestations. Conversely, when we meet, on one line of evolution, a recollection, so to speak, of what is developed along other lines, we must conclude that we have before us dissociated elements of one and the same original tendency. In this sense, vegetables and animals represent the two great divergent developments of life. Though the plant is distinguished from the animal by fixity and insensibility, movement and consciousness sleep in it as recollections which may waken. But, beside these normally sleeping recollections, there are others awake and active, just those, namely, whose activity does not obstruct the development of the elementary tendency itself. We may then formulate this law: _When a tendency splits up in the course of its development, each of the special tendencies which thus arise tries to preserve and develop everything in the primitive tendency that is not incompatible with the work for which it is specialized._ This explains precisely the fact we dwelt on in the preceding chapter, viz., the formation of identical complex mechanisms on independent lines of evolution. Certain deep-seated a.n.a.logies between the animal and the vegetable have probably no other cause: s.e.xual generation is perhaps only a luxury for the plant, but to the animal it was a necessity, and the plant must have been driven to it by the same impetus which impelled the animal thereto, a primitive, original impetus, anterior to the separation of the two kingdoms. The same may be said of the tendency of the vegetable towards a growing complexity. This tendency is essential to the animal kingdom, ever tormented by the need of more and more extended and effective action. But the vegetable, condemned to fixity and insensibility, exhibits the same tendency only because it received at the outset the same impulsion. Recent experiments show that it varies at random when the period of "mutation" arrives; whereas the animal must have evolved, we believe, in much more definite directions. But we will not dwell further on this original doubling of the modes of life. Let us come to the evolution of animals, in which we are more particularly interested.

What const.i.tutes animality, we said, is the faculty of utilizing a releasing mechanism for the conversion of as much stored-up potential energy as possible into "explosive" actions. In the beginning the explosion is haphazard, and does not choose its direction. Thus the amoeba thrusts out its pseudopodic prolongations in all directions at once. But, as we rise in the animal scale, the form of the body itself is observed to indicate a certain number of very definite directions along which the energy travels. These directions are marked by so many chains of nervous elements. Now, the nervous element has gradually emerged from the barely differentiated ma.s.s of organized tissue. It may, therefore, be surmised that in the nervous element, as soon as it appears, and also in its appendages, the faculty of suddenly freeing the gradually stored-up energy is concentrated. No doubt, every living cell expends energy without ceasing, in order to maintain its equilibrium.

The vegetable cell, torpid from the start, is entirely absorbed in this work of maintenance alone, as if it took for end what must at first have been only a means. But, in the animal, all points to action, that is, to the utilization of energy for movements from place to place. True, every animal cell expends a good deal--often the whole--of the energy at its disposal in keeping itself alive; but the organism as a whole tries to attract as much energy as possible to those points where the locomotive movements are effected. So that where a nervous system exists, with its complementary sense-organs and motor apparatus, everything should happen as if the rest of the body had, as its essential function, to prepare for these and pa.s.s on to them, at the moment required, that force which they are to liberate by a sort of explosion.

The part played by food amongst the higher animals is, indeed, extremely complex. In the first place it serves to repair tissues, then it provides the animal with the heat necessary to render it as independent as possible of changes in external temperature. Thus it preserves, supports, and maintains the organism in which the nervous system is set and on which the nervous elements have to live. But these nervous elements would have no reason for existence if the organism did not pa.s.s to them, and especially to the muscles they control, a certain energy to expend; and it may even be conjectured that there, in the main, is the essential and ultimate destination of food. This does not mean that the greater part of the food is used in this work. A state may have to make enormous expenditure to secure the return of taxes, and the sum which it will have to dispose of, after deducting the cost of collection, will perhaps be very small: that sum is, none the less, the reason for the tax and for all that has been spent to obtain its return.

So it is with the energy which the animal demands of its food.

Many facts seem to indicate that the nervous and muscular elements stand in this relation towards the rest of the organism. Glance first at the distribution of alimentary substances among the different elements of the living body. These substances fall into two cla.s.ses, one the quaternary or alb.u.minoid, the other the ternary, including the carbohydrates and the fats. The alb.u.minoids are properly plastic, destined to repair the tissues--although, owing to the carbon they contain, they are capable of providing energy on occasion. But the function of supplying energy has devolved more particularly on the second cla.s.s of substances: these, being deposited in the cell rather than forming part of its substance, convey to it, in the form of chemical potential, an expansive energy that may be directly converted into either movement or heat. In short, the chief function of the alb.u.minoids is to repair the machine, while the function of the other cla.s.s of substances is to supply power. It is natural that the alb.u.minoids should have no specially allotted destination, since every part of the machine has to be maintained. But not so with the other substances. The carbohydrates are distributed very unequally, and this inequality of distribution seems to us in the highest degree instructive.

Conveyed by the arterial blood in the form of glucose, these substances are deposited, in the form of glycogen, in the different cells forming the tissues. We know that one of the princ.i.p.al functions of the liver is to maintain at a constant level the quant.i.ty of glucose held by the blood, by means of the reserves of glycogen secreted by the hepatic cells. Now, in this circulation of glucose and acc.u.mulation of glycogen, it is easy to see that the effect is as if the whole effort of the organism were directed towards providing with potential energy the elements of both the muscular and the nervous tissues. The organism proceeds differently in the two cases, but it arrives at the same result. In the first case, it provides the muscle-cell with a large reserve deposited in advance: the quant.i.ty of glycogen contained in the muscles is, indeed, enormous in comparison with what is found in the other tissues. In the nervous tissue, on the contrary, the reserve is small (the nervous elements, whose function is merely to liberate the potential energy stored in the muscle, never have to furnish much work at one time); but the remarkable thing is that this reserve is restored by the blood at the very moment that it is expended, so that the nerve is instantly recharged with potential energy. Muscular tissue and nervous tissue are, therefore, both privileged, the one in that it is stocked with a large reserve of energy, the other in that it is always served at the instant it is in need and to the exact extent of its requirements.

More particularly, it is from the sensori-motor system that the call for glycogen, the potential energy, comes, as if the rest of the organism were simply there in order to transmit force to the nervous system and to the muscles which the nerves control. True, when we think of the part played by the nervous system (even the sensori-motor system) as regulator of the organic life, it may well be asked whether, in this exchange of good offices between it and the rest of the body, the nervous system is indeed a master that the body serves. But we shall already incline to this hypothesis when we consider, even in the static state only, the distribution of potential energy among the tissues; and we shall be entirely convinced of it when we reflect upon the conditions in which the energy is expended and restored. For suppose the sensori-motor system is a system like the others, of the same rank as the others. Borne by the whole of the organism, it will wait until an excess of chemical potential is supplied to it before it performs any work. In other words, it is the production of glycogen which will regulate the consumption by the nerves and muscles. On the contrary, if the sensori-motor system is the actual master, the duration and extent of its action will be independent, to a certain extent at least, of the reserve of glycogen that it holds, and even of that contained in the whole of the organism. It will perform work, and the other tissues will have to arrange as they can to supply it with potential energy. Now, this is precisely what does take place, as is shown in particular by the experiments of Morat and Dufourt.[56] While the glycogenic function of the liver depends on the action of the excitory nerves which control it, the action of these nerves is subordinated to the action of those which stimulate the locomotor muscles--in this sense, that the muscles begin by expending without calculation, thus consuming glycogen, impoveris.h.i.+ng the blood of its glucose, and finally causing the liver, which has had to pour into the impoverished blood some of its reserve of glycogen, to manufacture a fresh supply. From the sensori-motor system, then, everything starts; on that system everything converges; and we may say, without metaphor, that the rest of the organism is at its service.

Consider again what happens in a prolonged fast. It is a remarkable fact that in animals that have died of hunger the brain is found to be almost unimpaired, while the other organs have lost more or less of their weight and their cells have undergone profound changes.[57] It seems as though the rest of the body had sustained the nervous system to the last extremity, treating itself simply as the means of which the nervous system is the end.

To sum up: if we agree, in short, to understand by "the sensori-motor system" the cerebro-spinal nervous system together with the sensorial apparatus in which it is prolonged and the locomotor muscles it controls, we may say that a higher organism is essentially a sensori-motor system installed on systems of digestion, respiration, circulation, secretion, etc., whose function it is to repair, cleanse and protect it, to create an unvarying internal environment for it, and above all to pa.s.s it potential energy to convert into locomotive movement.[58] It is true that the more the nervous function is perfected, the more must the functions required to maintain it develop, and the more exacting, consequently, they become for themselves. As the nervous activity has emerged from the protoplasmic ma.s.s in which it was almost drowned, it has had to summon around itself activities of all kinds for its support. These could only be developed on other activities, which again implied others, and so on indefinitely. Thus it is that the complexity of functioning of the higher organisms goes on to infinity. The study of one of these organisms therefore takes us round in a circle, as if everything was a means to everything else. But the circle has a centre, none the less, and that is the system of nervous elements stretching between the sensory organs and the motor apparatus.

We will not dwell here on a point we have treated at length in a former work. Let us merely recall that the progress of the nervous system has been effected both in the direction of a more precise adaptation of movements and in that of a greater lat.i.tude left to the living being to choose between them. These two tendencies may appear antagonistic, and indeed they are so; but a nervous chain, even in its most rudimentary form, successfully reconciles them. On the one hand, it marks a well-defined track between one point of the periphery and another, the one sensory, the other motor. It has therefore ca.n.a.lized an activity which was originally diffused in the protoplasmic ma.s.s. But, on the other hand, the elements that compose it are probably discontinuous; at any rate, even supposing they anastomose, they exhibit a _functional_ discontinuity, for each of them ends in a kind of cross-road where probably the nervous current may choose its course. From the humblest Monera to the best endowed insects, and up to the most intelligent vertebrates, the progress realized has been above all a progress of the nervous system, coupled at every stage with all the new constructions and complications of mechanism that this progress required. As we foreshadowed in the beginning of this work, the role of life is to insert some _indetermination_ into matter. Indeterminate, _i.e._ unforeseeable, are the forms it creates in the course of its evolution.

More and more indeterminate also, more and more free, is the activity to which these forms serve as the vehicle. A nervous system, with neurones placed end to end in such wise that, at the extremity of each, manifold ways open in which manifold questions present themselves, is a veritable _reservoir of indetermination_. That the main energy of the vital impulse has been spent in creating apparatus of this kind is, we believe, what a glance over the organized world as a whole easily shows.

But concerning the vital impulse itself a few explanations are necessary.

It must not be forgotten that the force which is evolving throughout the organized world is a limited force, which is always seeking to transcend itself and always remains inadequate to the work it would fain produce.

The errors and puerilities of radical finalism are due to the misapprehension of this point. It has represented the whole of the living world as a construction, and a construction a.n.a.logous to a human work. All the pieces have been arranged with a view to the best possible functioning of the machine. Each species has its reason for existence, its part to play, its allotted place; and all join together, as it were, in a musical concert, wherein the seeming discords are really meant to bring out a fundamental harmony. In short, all goes on in nature as in the works of human genius, where, though the result may be trifling, there is at least perfect adequacy between the object made and the work of making it.

Nothing of the kind in the evolution of life. There, the disproportion is striking between the work and the result. From the bottom to the top of the organized world we do indeed find one great effort; but most often this effort turns short, sometimes paralyzed by contrary forces, sometimes diverted from what it should do by what it does, absorbed by the form it is engaged in taking, hypnotized by it as by a mirror. Even in its most perfect works, though it seems to have triumphed over external resistances and also over its own, it is at the mercy of the materiality which it has had to a.s.sume. It is what each of us may experience in himself. Our freedom, in the very movements by which it is affirmed, creates the growing habits that will stifle it if it fails to renew itself by a constant effort: it is dogged by automatism. The most living thought becomes frigid in the formula that expresses it. The word turns against the idea.

The letter kills the spirit. And our most ardent enthusiasm, as soon as it is externalized into action, is so naturally congealed into the cold calculation of interest or vanity, the one takes so easily the shape of the other, that we might confuse them together, doubt our own sincerity, deny goodness and love, if we did not know that the dead retain for a time the features of the living.

The profound cause of this discordance lies in an irremediable difference of rhythm. Life in general is mobility itself; particular manifestations of life accept this mobility reluctantly, and constantly lag behind. It is always going ahead; they want to mark time. Evolution in general would fain go on in a straight line; each special evolution is a kind of circle. Like eddies of dust raised by the wind as it pa.s.ses, the living turn upon themselves, borne up by the great blast of life. They are therefore relatively stable, and counterfeit immobility so well that we treat each of them as a _thing_ rather than as a _progress_, forgetting that the very permanence of their form is only the outline of a movement. At times, however, in a fleeting vision, the invisible breath that bears them is materialized before our eyes. We have this sudden illumination before certain forms of maternal love, so striking, and in most animals so touching, observable even in the solicitude of the plant for its seed. This love, in which some have seen the great mystery of life, may possibly deliver us life's secret. It shows us each generation leaning over the generation that shall follow.

It allows us a glimpse of the fact that the living being is above all a thoroughfare, and that the essence of life is in the movement by which life is transmitted.

This contrast between life in general, and the forms in which it is manifested, has everywhere the same character. It might be said that life tends toward the utmost possible action, but that each species prefers to contribute the slightest possible effort. Regarded in what const.i.tutes its true essence, namely, as a transition from species to species, life is a continually growing action. But each of the species, through which life pa.s.ses, aims only at its own convenience. It goes for that which demands the least labor. Absorbed in the form it is about to take, it falls into a partial sleep, in which it ignores almost all the rest of life; it fas.h.i.+ons itself so as to take the greatest possible advantage of its immediate environment with the least possible trouble.

Accordingly, the act by which life goes forward to the creation of a new form, and the act by which this form is shaped, are two different and often antagonistic movements. The first is continuous with the second, but cannot continue in it without being drawn aside from its direction, as would happen to a man leaping, if, in order to clear the obstacle, he had to turn his eyes from it and look at himself all the while.

Living forms are, by their very definition, forms that are able to live.

In whatever way the adaptation of the organism to its circ.u.mstances is explained, it has necessarily been sufficient, since the species has subsisted. In this sense, each of the successive species that paleontology and zoology describes was a _success_ carried off by life.

But we get a very different impression when we refer each species to the movement that has left it behind on its way, instead of to the conditions into which it has been set. Often this movement has turned aside; very often, too, it has stopped short; what was to have been a thoroughfare has become a terminus. From this new point of view, failure seems the rule, success exceptional and always imperfect. We shall see that, of the four main directions along which animal life bent its course, two have led to blind alleys, and, in the other two, the effort has generally been out of proportion to the result.

Doc.u.ments are lacking to reconstruct this history in detail, but we can make out its main lines. We have already said that animals and vegetables must have separated soon from their common stock, the vegetable falling asleep in immobility, the animal, on the contrary, becoming more and more awake and marching on to the conquest of a nervous system. Probably the effort of the animal kingdom resulted in creating organisms still very simple, but endowed with a certain freedom of action, and, above all, with a shape so undecided that it could lend itself to any future determination. These animals may have resembled some of our worms, but with this difference, however, that the worms living to-day, to which they could be compared, are but the empty and fixed examples of infinitely plastic forms, pregnant with an unlimited future, the common stock of the echinoderms, molluscs, arthropods, and vertebrates.

One danger lay in wait for them, one obstacle which might have stopped the soaring course of animal life. There is one peculiarity with which we cannot help being struck when glancing over the fauna of primitive times, namely, the imprisonment of the animal in a more or less solid sheath, which must have obstructed and often even paralyzed its movements. The molluscs of that time had a sh.e.l.l more universally than those of to-day. The arthropods in general were provided with a carapace; most of them were crustaceans. The more ancient fishes had a bony sheath of extreme hardness.[59] The explanation of this general fact should be sought, we believe, in a tendency of soft organisms to defend themselves against one another by making themselves, as far as possible, undevourable. Each species, in the act by which it comes into being, trends towards that which is most expedient. Just as among primitive organisms there were some that turned towards animal life by refusing to manufacture organic out of inorganic material and taking organic substances ready made from organisms that had turned toward the vegetative life, so, among the animal species themselves, many contrived to live at the expense of other animals. For an organism that is animal, that is to say mobile, can avail itself of its mobility to go in search of defenseless animals, and feed on them quite as well as on vegetables.

So, the more species became mobile, the more they became voracious and dangerous to one another. Hence a sudden arrest of the entire animal world in its progress towards higher and higher mobility; for the hard and calcareous skin of the echinoderm, the sh.e.l.l of the mollusc, the carapace of the crustacean and the ganoid breast-plate of the ancient fishes probably all originated in a common effort of the animal species to protect themselves against hostile species. But this breast-plate, behind which the animal took shelter, constrained it in its movements and sometimes fixed it in one place. If the vegetable renounced consciousness in wrapping itself in a cellulose membrane, the animal that shut itself up in a citadel or in armor condemned itself to a partial slumber. In this torpor the echinoderms and even the molluscs live to-day. Probably arthropods and vertebrates were threatened with it too. They escaped, however, and to this fortunate circ.u.mstance is due the expansion of the highest forms of life.

In two directions, in fact, we see the impulse of life to movement getting the upper hand again. The fishes exchanged their ganoid breast-plate for scales. Long before that, the insects had appeared, also disenc.u.mbered of the breast-plate that had protected their ancestors. Both supplemented the insufficiency of their protective covering by an agility that enabled them to escape their enemies, and also to a.s.sume the offensive, to choose the place and the moment of encounter. We see a progress of the same kind in the evolution of human armaments. The first impulse is to seek shelter; the second, which is the better, is to become as supple as possible for flight and above all for attack--attack being the most effective means of defense. So the heavy hoplite was supplanted by the legionary; the knight, clad in armor, had to give place to the light free-moving infantryman; and in a general way, in the evolution of life, just as in the evolution of human societies and of individual destinies, the greatest successes have been for those who have accepted the heaviest risks.

Evidently, then, it was to the animal's interest to make itself more mobile. As we said when speaking of adaptation in general, any transformation of a species can be explained by its own particular interest. This will give the immediate cause of the variation, but often only the most superficial cause. The profound cause is the impulse which thrust life into the world, which made it divide into vegetables and animals, which shunted the animal on to suppleness of form, and which, at a certain moment, in the animal kingdom threatened with torpor, secured that, on some points at least, it should rouse itself up and move forward.

On the two paths along which the vertebrates and arthropods have separately evolved, development (apart from retrogressions connected with parasitism or any other cause) has consisted above all in the progress of the sensori-motor nervous system. Mobility and suppleness were sought for, and also--through many experimental attempts, and not without a tendency to excess of substance and brute force at the start--variety of movements. But this quest itself took place in divergent directions. A glance at the nervous system of the arthropods and that of the vertebrates shows us the difference. In the arthropods, the body is formed of a series more or less long of rings set together; motor activity is thus distributed amongst a varying--sometimes a considerable--number of appendages, each of which has its special function. In the vertebrates, activity is concentrated in two pairs of members only, and these organs perform functions which depend much less strictly on their form.[60] The independence becomes complete in man, whose hand is capable of any kind of work.

That, at least, is what we see. But behind what is seen there is what may be surmised--two powers, immanent in life and originally intermingled, which were bound to part company in course of growth.

To define these powers, we must consider, in the evolution both of the arthropods and the vertebrates, the species which mark the culminating point of each. How is this point to be determined? Here again, to aim at geometrical precision will lead us astray. There is no single simple sign by which we can recognize that one species is more advanced than another on the same line of evolution. There are manifold characters, that must be compared and weighed in each particular case, in order to ascertain to what extent they are essential or accidental and how far they must be taken into account.

It is unquestionable, for example, that _success_ is the most general criterion of superiority, the two terms being, up to a certain point, synonymous. By success must be understood, so far as the living being is concerned, an apt.i.tude to develop in the most diverse environments, through the greatest possible variety of obstacles, so as to cover the widest possible extent of ground. A species which claims the entire earth for its domain is truly a dominating and consequently superior species. Such is the human species, which represents the culminating point of the evolution of the vertebrates. But such also are, in the series of the articulate, the insects and in particular certain hymenoptera. It has been said of the ants that, as man is lord of the soil, they are lords of the sub-soil.

On the other hand, a group of species that has appeared late may be a group of degenerates; but, for that, some special cause of retrogression must have intervened. By right, this group should be superior to the group from which it is derived, since it would correspond to a more advanced stage of evolution. Now man is probably the latest comer of the vertebrates;[61] and in the insect series no species is later than the hymenoptera, unless it be the lepidoptera, which are probably degenerates, living parasitically on flowering plants.

So, by different ways, we are led to the same conclusion. The evolution of the arthropods reaches its culminating point in the insect, and in particular in the hymenoptera, as that of the vertebrates in man. Now, since instinct is nowhere so developed as in the insect world, and in no group of insects so marvelously as in the hymenoptera, it may be said that the whole evolution of the animal kingdom, apart from retrogressions towards vegetative life, has taken place on two divergent paths, one of which led to instinct and the other to intelligence.

Vegetative torpor, instinct, and intelligence--these, then, are the elements that coincided in the vital impulsion common to plants and animals, and which, in the course of a development in which they were made manifest in the most unforeseen forms, have been dissociated by the very fact of their growth. _The cardinal error which, from Aristotle onwards, has vitiated most of the philosophies of nature, is to see in vegetative, instinctive and rational life, three successive degrees of the development of one and the same tendency, whereas they are three divergent directions of an activity that has split up as it grew._ The difference between them is not a difference of intensity, nor, more generally, of degree, but of kind.

It is important to investigate this point. We have seen in the case of vegetable and animal life how they are at once mutually complementary and mutually antagonistic. Now we must show that intelligence and instinct also are opposite and complementary. But let us first explain why we are generally led to regard them as activities of which one is superior to the other and based upon it, whereas in reality they are not things of the same order: they have not succeeded one another, nor can we a.s.sign to them different grades.

It is because intelligence and instinct, having originally been interpenetrating, retain something of their common origin. Neither is ever found in a pure state. We said that in the plant the consciousness and mobility of the animal, which lie dormant, can be awakened; and that the animal lives under the constant menace of being drawn aside to the vegetative life. The two tendencies--that of the plant and that of the animal--were so thoroughly interpenetrating, to begin with, that there has never been a complete severance between them: they haunt each other continually; everywhere we find them mingled; it is the proportion that differs. So with intelligence and instinct. There is no intelligence in which some traces of instinct are not to be discovered, more especially no instinct that is not surrounded with a fringe of intelligence. It is this fringe of intelligence that has been the cause of so many misunderstandings. From the fact that instinct is always more or less intelligent, it has been concluded that instinct and intelligence are things of the same kind, that there is only a difference of complexity or perfection between them, and, above all, that one of the two is expressible in terms of the other. In reality, they accompany each other only because they are complementary, and they are complementary only because they are different, what is instinctive in instinct being opposite to what is intelligent in intelligence.

We are bound to dwell on this point. It is one of the utmost importance.

Let us say at the outset that the distinctions we are going to make will be too sharply drawn, just because we wish to define in instinct what is instinctive, and in intelligence what is intelligent, whereas all concrete instinct is mingled with intelligence, as all real intelligence is penetrated by instinct. Moreover, neither intelligence nor instinct lends itself to rigid definition: they are tendencies, and not things.

Also, it must not be forgotten that in the present chapter we are considering intelligence and instinct as going out of life which deposits them along its course. Now the life manifested by an organism is, in our view, a certain effort to obtain certain things from the material world. No wonder, therefore, if it is the diversity of this effort that strikes us in instinct and intelligence, and if we see in these two modes of psychical activity, above all else, two different methods of action on inert matter. This rather narrow view of them has the advantage of giving us an objective means of distinguis.h.i.+ng them. In return, however, it gives us, of intelligence in general and of instinct in general, only the mean position above and below which both constantly oscillate. For that reason the reader must expect to see in what follows only a diagrammatic drawing, in which the respective outlines of intelligence and instinct are sharper than they should be, and in which the shading-off which comes from the indecision of each and from their reciprocal encroachment on one another is neglected. In a matter so obscure, we cannot strive too hard for clearness. It will always be easy afterwards to soften the outlines and to correct what is too geometrical in the drawing--in short, to replace the rigidity of a diagram by the suppleness of life.

To what date is it agreed to ascribe the appearance of man on the earth?

To the period when the first weapons, the first tools, were made. The memorable quarrel over the discovery of Boucher de Perthes in the quarry of Moulin-Quignon is not forgotten. The question was whether real hatchets had been found or merely bits of flint accidentally broken. But that, supposing they were hatchets, we were indeed in the presence of intelligence, and more particularly of _human_ intelligence, no one doubted for an instant. Now let us open a collection of anecdotes on the intelligence of animals: we shall see that besides many acts explicable by imitation or by the automatic a.s.sociation of images, there are some that we do not hesitate to call intelligent: foremost among them are those that bear witness to some idea of manufacture, whether the animal life succeeds in fas.h.i.+oning a crude instrument or uses for its profit an object made by man. The animals that rank immediately after man in the matter of intelligence, the apes and elephants, are those that can use an artificial instrument occasionally. Below, but not very far from them, come those that _recognize_ a constructed object: for example, the fox, which knows quite well that a trap is a trap. No doubt, there is intelligence wherever there is inference; but inference, which consists in an inflection of past experience in the direction of present experience, is already a beginning of invention. Invention becomes complete when it is materialized in a manufactured instrument. Towards that achievement the intelligence of animals tends as towards an ideal.

And though, ordinarily, it does not yet succeed in fas.h.i.+oning artificial objects and in making use of them, it is preparing for this by the very variations which it performs on the instincts furnished by nature. As regards human intelligence, it has not been sufficiently noted that mechanical invention has been from the first its essential feature, that even to-day our social life gravitates around the manufacture and use of artificial instruments, that the inventions which strew the road of progress have also traced its direction. This we hardly realize, because it takes us longer to change ourselves than to change our tools. Our individual and even social habits survive a good while the circ.u.mstances for which they were made, so that the ultimate effects of an invention are not observed until its novelty is already out of sight. A century has elapsed since the invention of the steam-engine, and we are only just beginning to feel the depths of the shock it gave us. But the revolution it has effected in industry has nevertheless upset human relations altogether. New ideas are arising, new feelings are on the way to flower. In thousands of years, when, seen from the distance, only the broad lines of the present age will still be visible, our wars and our revolutions will count for little, even supposing they are remembered at all; but the steam-engine, and the procession of inventions of every kind that accompanied it, will perhaps be spoken of as we speak of the bronze or of the chipped stone of prehistoric times: it will serve to define an age.[62] If we could rid ourselves of all pride, if, to define our species, we kept strictly to what the historic and the prehistoric periods show us to be the constant characteristic of man and of intelligence, we should say not _h.o.m.o sapiens_, but _h.o.m.o faber_. In short, _intelligence, considered in what seems to be its original feature, is the faculty of manufacturing artificial objects, especially tools to make tools, and of indefinitely varying the manufacture_.