On the Genesis of Species Part 14

It will, however, be replied that such an ent.i.ty is no _vera causa_; that if the conception is accepted, it is no real explanation; and that it is merely a roundabout way of saying that the facts are as they are, while the cause remains unknown. To this it may be rejoined that for all who believe in the existence of the abstraction "force" at all, other than will, {228} this conception of an internal force must be accepted and located somewhere--cannot be eliminated altogether; and that therefore it may as reasonably be accepted in this mode as in any other.

It was urged at the end of the third chapter that it is congruous to credit mineral species with an internal power or force. By such a power it may be conceived that crystals not only a.s.sume their external symmetry, but even repair it when injured. Ultimate chemical elements must also be conceived as possessing an innate tendency to form certain unions, and to cohere in stable aggregations. This was considered towards the end of Chapter VIII.

Turning to the organic world, even on the hypothesis of Mr. Herbert Spencer or that of Mr. Darwin, it is impossible to escape the conception of innate internal forces. With regard to the physiological units of the former, Mr.

Spencer himself, as we have seen, distinctly attributes to them "an _innate_ tendency" to evolve the parent form from which they sprang. With regard to the gemmules of Mr. Darwin, we have seen, in Chapter X., with how many innate powers, tendencies, and capabilities they must each be severally endowed, to reproduce their kind, to evolve complex organisms or cells, to exercise germinative affinity, &c.

If then (as was before said at the end of Chapter VIII.) such innate powers must be attributed to chemical atoms, to mineral species, to gemmules, and to physiological units, it is only reasonable to attribute such to each individual organism.

The conception of such internal and latent capabilities is somewhat like that of Mr. Galton, before mentioned, according to which the organic world consists of ent.i.ties, each of which is, as it were, a spheroid with many facets on its surface, upon one of which it reposes in stable equilibrium.

When by the acc.u.mulated action of incident forces this equilibrium is {229} disturbed, the spheroid is supposed to turn over until it settles on an adjacent facet once more in stable equilibrium.

The internal tendency of an organism to certain considerable and definite changes would correspond to the facets on the surface of the spheroid.

It may be objected that we have no knowledge as to how terrestrial, cosmical and other forces can affect organisms so as to stimulate and evolve these latent, merely potential forms. But we have had evidence that such mysterious agencies _do_ affect organisms in ways as yet inexplicable, in the very remarkable effects of geographical conditions which were detailed in the third chapter.

It is quite conceivable that the material organic world may be so const.i.tuted that the simultaneous action upon it of all known forces, mechanical, physical, chemical, magnetic, terrestrial, and cosmical, together with other as yet unknown forces which probably exist, may result in changes which are harmonious and symmetrical, just as the internal nature of vibrating plates causes particles of sand scattered over them to a.s.sume definite and symmetrical figures when made to oscillate in different ways by the bow of a violin being drawn along their edges. The results of these combined internal powers and external influences might be represented under the symbol of complex series of vibrations (a.n.a.logous to those of sound or light) forming a most complex harmony or a display of most varied colours. In such a way the reparation of local injuries might be symbolized as a filling up and completion of an interrupted rhythm. Thus also monstrous aberrations from typical structure might correspond to a discord, and sterility from crossing be compared with the darkness resulting from the interference of waves of light.

Such symbolism will harmonize with the peculiar reproduction, before mentioned, of heads in the body of certain annelids, with the facts of serial h.o.m.ology, as well as those of bilateral and vertical symmetry. {230} Also, as the atoms of a resonant body may be made to give out sound by the juxtaposition of a vibrating tuning-fork, so it is conceivable that the physiological units of a living organism may be so influenced by surrounding conditions (organic and other) that the acc.u.mulation of these conditions may upset the previous rhythm of such units, producing modifications in them--a fresh chord in the harmony of nature--a new species!

But it may be again objected that to say that species arise by the help of an innate power possessed by organisms is no explanation, but is a reproduction of the absurdity, _l'opium endormit parcequ'il a une vertu soporifique_. It is contended, however, that this objection does not apply, even if it be conceded that there is that force in Moliere's ridicule which is generally attributed to it.[231] Much, however, might be said in opposition to more than one of that brilliant dramatist's smart philosophical epigrams, just as to the theological ones of Voltaire, or to the biological one of that other Frenchman who for a time discredited a cranial skeletal theory by the phrase "Vertebre pensante."[232]

In fact, however, it is a real explanation of how a man lives to say that he lives independently, on his own income, instead of being supported by his relatives and friends. In the same way, there is fully as real a distinction between the production of new specific manifestations entirely _ab externo_, and by the production of the same through an innate force and tendency, the determination of which into action is occasioned by {231} external circ.u.mstances.

To say that organisms possess this innate power, and that by it new species are from time to time produced, is by no means a mere a.s.sertion that they _are_ produced, and in an unknown mode. It is the negation of that view which deems external forces alone sufficient, and at the same time the a.s.sertion of something positive, to be arrived at by the process of _reductio ad absurdum_.

All physical explanations result ultimately in such conceptions of innate power, or else in that of will force. The far-famed explanation of the celestial motions ends in the conception that every particle of matter has the innate power of attracting every other particle directly as the ma.s.s, and inversely as the square of the distance.

We are logically driven to this positive conception if we do not accept the view that there is no force but volition, and that all phenomena whatever are the immediate results of the action of intelligent and self-conscious will.

We have seen that the notion of sudden changes--saltatory actions in nature--has received countenance from Professor Huxley.[233] We must conceive that these jumps are orderly, and according to law, inasmuch as the whole cosmos is such. Such orderly evolution harmonizes with a teleology derived, not indeed from external nature directly, but from the mind of man. On this point, however, more will be said in the next chapter.

But, once more, if new species are not manifested by the action of external conditions upon minute indefinite individual differences, in what precise way may we conceive that manifestation to have taken place?

Are new species now evolving, as they have been from time to time evolved?

If so, in what way and by what conceivable means?

{232} In the first place, they must be produced by natural action in pre-existing material, or by supernatural action.

For reasons to be given in the next chapter, the second hypothesis need not be considered.

If, then, new species are and have been evolved from pre-existing material, must that material have been organic or inorganic?

As before said, additional arguments have lately been brought forward to show that individual organisms _do_ arise from a basis of _in_-organic material only. As, however, this at the most appears to be the case, if at all, only with the lowest and most minute organisms exclusively, the process cannot be observed, though it may perhaps be fairly inferred.

We may therefore, if for no other reason, dismiss the notion that highly organized animals and plants can be suddenly or gradually built up by any combination of physical forces and natural powers acting externally and internally upon and in merely inorganic material as a base.

But the question is, how have the highest kinds of animals and plants arisen? It seems impossible that they can have appeared otherwise than by the agency of antecedent organisms not greatly different from them.

A mult.i.tude of facts, ever increasing in number and importance, all point to such a mode of specific manifestation.

One very good example has been adduced by Professor Flower in the introductory lecture of his first Hunterian Course.[234] It is the reduction in size, to a greater or less degree, of the second and third digits of the foot in Australian marsupials, and this, in spite of the very different form and function of the foot in different groups of those animals.

A similarly significant evidence of relations.h.i.+p is afforded by processes of the zygomatic region of the skull in certain edentates existing and extinct.

{233} Again, the relation between existing and recent faunas of the different regions of the world, and the predominating (though by no means exclusive) march of organization, from the more general to the more special, point in the same direction.

Almost all the facts brought forward by the patient industry of Mr. Darwin in support of his theory of "Natural Selection," are of course available as evidence in favour of the agency of pre-existing and similar animals in specific evolution.

Now the new forms must be produced by changes taking place in organisms in, after or before their birth, either in their embryonic, or towards or in their adult, condition.

Examples of strange births are sufficiently common, and they may arise either from direct embryonic modifications or apparently from some obscure change in the parental action. To the former category belong the hosts of instances of malformation through arrest of development, and perhaps generally monstrosities of some sort are the result of such affections of the embryo. To the second category belong all cases of hybridism, of cross breed, and in all probability the new varieties and forms, such as the memorable one of the black-shouldered peac.o.c.k. In all these cases we do not have abortions or monstrosities, but more or less harmonious forms often of great functional activity, endowed with marked viability and generative prepotency, except in the case of hybrids, when we often find even a more marked generative impotency.

It seems probable therefore that new species may arise from some const.i.tutional affection of parental forms--an affection mainly, if not exclusively, of their generative system. Mr. Darwin has carefully collected[235] numerous instances to show how excessively sensitive to various influences this system is. He says:[236] "Sterility is independent of general health, and is often accompanied by excess of size, or {234} great luxuriance," and, "No one can tell, till he tries; whether any particular animal will breed under confinement, or any exotic plant seed freely under culture." Again, "When a new character arises, whatever its nature may be, it generally tends to be inherited, at least in a temporary and sometimes in a most persistent manner."[237] Yet the obscure action of conditions will alter characters long inherited, as the grandchildren of Aylesbury ducks, removed to a distant part of England, completely lost their early habit of incubation, and hatched their eggs at the same time with the common ducks of the same place.[238]

Mr. Darwin quotes Mr. Bartlett as saying: "It is remarkable that lions breed more freely in travelling collections than in the zoological gardens; probably the constant excitement and irritation produced by moving from place to place, or change of air, may have considerable influence in the matter."[239]

Mr. Darwin also says: "There is reason to believe that insects are affected by confinement like the higher animals," and he gives examples.[240]

Again, he gives examples of change of plumage in the linnet, bunting, oriole, and other birds, and of the temporary modification of the horns of a male deer during a voyage.[241]

Finally, he adds that these changes cannot be attributed to loss of health or vigour, "when we reflect how healthy, long-lived, and vigorous many animals are under captivity, such as parrots, and hawks when used for hawking, chetahs when used for hunting, and elephants. The reproductive organs themselves are not diseased; and the diseases from which animals in menageries usually perish, are not those which in any way affect their fertility. No domestic animal is more subject to disease than the sheep, yet it is remarkably prolific.... It would appear that any change in {235} the habits of life, whatever these habits may be, if great enough, tends to affect in an inexplicable manner the powers of reproduction."

Such, then, is the singular sensitiveness of the generative system.

As to the means by which that system is affected, we see that a variety of conditions affect it; but as to the modes in which they act upon it, we have as yet little if any clue.

We have also seen the singular effects (in tailed Lepidoptera, &c.) of causes connected with geographical distribution, the mode of action of which is as yet quite inexplicable; and we have also seen the innate tendency which there appears to be in certain groups (birds of paradise, &c.) to develop peculiarities of a special kind.

It is, to say the least, probable that other influences exist, terrestrial and cosmical, as yet un-noted. The gradually acc.u.mulating or diversely combining actions of all these on highly sensitive structures, which are themselves possessed of internal responsive powers and tendencies, may well result in occasional repeated productions of forms harmonious and vigorous, and differing from the parental forms in proportion to the result of the combining or conflicting action of all external and internal influences.

If, in the past history of this planet, more causes ever intervened, or intervened more energetically than at present, we might _a priori_ expect a richer and more various evolution of forms more radically differing than any which could be produced under conditions of more perfect equilibrium.

At the same time, if it be true that the last few thousand years have been a period of remarkable and exceptional uniformity as regards this planet's astronomical relations, there are then some grounds for thinking that organic evolution may have been exceptionally depressed during the same epoch.

Now, as to the fact that sudden changes and sudden developments have {236} occurred, and as to the probability that such changes are likely to occur, evidence was given in Chapter IV.

In Chapter V. we also saw that minerals become modified suddenly and considerably by the action of incident forces--as, _e.g._, the production of hexagonal tabular crystals of carbonate of copper by sulphuric acid, and of long rectangular prisms by ammonia, &c.

We have thus a certain antecedent probability that if changes are produced in specific manifestation through incident forces, these changes will be sensible and considerable, not minute and infinitesimal.

Consequently, it is probable that new species have appeared from time to time with comparative suddenness, and that they still continue so to arise if all the conditions necessary for specific evolution now obtain.

This probability will be increased if the observations of Dr. Bastian are confirmed by future investigation. According to his report, when the requisite conditions were supplied, the transformations which appeared to take place (from very low to higher organisms) were sudden, definite, and complete.

Therefore, if this is so, there must probably exist in higher forms a similar tendency to such change. That tendency may indeed be long suppressed, and ultimately modified by the action of heredity--an action which would increase in force with the increase in the perfection and complexity of the organism affected. Still we might expect that such changes as do take place would be also sudden, definite, and complete.

Moreover, as the same causes produce the same effects, several individual parent forms must often have been similarly and simultaneously affected.

That they should be so affected--at least that several similarly modified individuals should simultaneously arise--has been seen to be a generally necessary circ.u.mstance for the permanent duration of such new modifications.

It is also conceivable that such new forms may be endowed with {237} excessive const.i.tutional strength and viability, and with generative prepotency, as was the case with the black-shouldered peac.o.c.k in Sir J.